I don’t think there’s a problem here, it’s just a setup issue. I think all of the pieces to an answer are here, but let me see if I can bring them all together for you.
Mach 4 (like a lot of other control programs) allows for multiple “work offsets”. They are numbered G54, G55, G56, etc. You saw those in your “offsets” tab.
Think of these like saved points on a map, you can have a lot of different saved places. You can set these manually by typing numbers in, by driving your machine to a particular location and zeroing there, or by using the touch plate.
So let’s say you’re on the G54 offset and you zero to a piece of wood you have on the table. The DRO will say XYZ 0, but if you look in the offset table you’ll see numbers for XYZ. That’s because these are offsets from the MACHINE zero. Essentially these numbers are a distance off of your home switches (Z will be negative because Z zero is at the top)
As you may have gathered you can have multiple work offsets, but to keep matters simple lets just focus on having one, the G54 one. Before you zero you need to make sure it’s selected in the offsets tab.
Now let’s move on to Fusion. When you setup a job in Fusion you have to setup your stock (as seen in the video posted above). When doing that you have to setup a WCS zero point. As noted in the video Fusion often picks the center top of the stock as default. This is likely NOT what you want.
If you used your zero plate to set your G54 offset to the top left of your material you’re going to cut, than the WCS in Fusion needs to be in the same spot.
If you left it at the center of your stock the whole cut will be shifted in two directions half the size of your stock.
Now the last thing you need to make sure of is that when you post your G Code from Fusion make sure it’s set to the work offset you’re using, in this case G54 How to define Work Coordinate Systems in Fusion 360 Manufacture | Fusion 360 | Autodesk Knowledge Network
I think what was happening to you is at some point you picked a different WCS in Mach, and perhaps even switched to machine coordinates and you got confused with the numbers there. Things likely got more confusing when you perhaps didn’t have the WCS in the Fusion stock setup perfect, and to add even MORE confusion you may have changed the WCS that Fusion used in post process. Basically there three places you could have made an accidental wrong choice.
Hope that helps, report back and let us know!